- McComb, D., and G. F. Knowlton. 1953. Utah records of insect enemies of western forests. Utah Agricultural Experiment Station, Mimeograph Series 398. 5 p.
- Knowlton, G. F. 1931. Notes on Utah Coleoptera . Florida Entomologist 15:10.
- Knowlton, G. F., and G. P. Taylor. 1952. Beetles—Coleoptera: records and notes, largely from Utah . Utah Agricultural Experiment Station, Mimeograph Series 389:1-32.