- Smith, S. G., and N. Virkki. 1978. Animal cytogenetics. 3. Insecta 5. Coleoptera. Gebruder Borntrager, Berlin. 366 p.
- Virkki, N. 1960. Cytology of male meiosis in certain European forest beetles of the families Scolytidae, Cleridae, and Anobiidae. Annales Academiae Scientiarum Fennicae, Ser. A, IV. Biol. 49:3-16.