- Furniss, M. M., B. H. Baker, R. A. Werner, and L. C. Yarger. 1979. Characteristics of spruce beetle (Coleoptera) infestation in felled white spruce in Alaska . Canadian Entomologist 111:1355-1360.
- Graham, K., and A. E. Werner. 1956a. Chemical aspects of log selection by ambrosia beetles . Canada Department of Agriculture, Science Service, Forest Biology Division, Bi-monthly Progress Report 12:3-4.
- Graham, K., and A. E. Werner. 1956b. Chemical aspects of log selection by ambrosia beetles. Interim report 1955-1, Forest Biology Laboratory, Victoria, British Columbia. Canada Department of Agriculture, Science Service, Forest Biology Division. 25 p.
- Hard, J. S., R. A. Werner, and E. H. Holsten. 1983. Susceptibility of white spruce (Picea glauca) to attack by spruce beetles (Dendroctonus rufipennis) during the early years of an outbreak in Alaska . Canadian Journal of Forest Research 13:678-684.
- Holsten, E. H., and R. A. Werner. 1984. Evaluation of methyocyclohexenone (MCH) in preventing or suppressing spruce beetle attacks in Alaska. United States Department of Agriculture, Forest Service, Alaska Region, State and Private Forestry, Technical Report R-10-6. 16 p.
- Holsten, E. H., R. A. Werner, and T. H. Laurent. 1980. Insects and/diseases of Alaskan forests. United States Department of Agriculture, Forest Service, Alaska Region, Report 75. 187 p.
- Werner, A. E., and K. Graham. 1957. Volatile wood constituents in relation to ambrosia beetles . Canada Department of Agriculture, Science Service, Division of Forest Biology, Bi-monthly Progress Report 13:3.
- Werner, R. A. 1970. Laboratory studies on the olfactory response of Ips grandicollis (Eichh.) (Coleoptera: Scolytidae) to chemical attractants. Unpublished dissertation, North Carolina State University, Raleigh. 159 p.
- Werner, R. A. 1971. Studies on the olfactory response of Ips grandicollis (Eichhoff) (Coleoptera: Scolytidae) to host- and insect-produced attractants. Dissertation Abstracts 32(3):1639-B.
- Werner, R. A. 1972a. Aggregation behavior of the beetle Ips grandicollis in response to host produced attractants . Journal of Insect Physiology 18:423-437.
- Werner, R. A. 1972b. Aggregation behavior of the beetle Ips grandicollis in response to insect-produced attractants . Journal of Insect Physiology 18:1001-1013.
- Werner, R. A. 1972c. Response of the beetle Ips grandicollis to combinations of host and insect produced attractants . Journal of Insect Physiology 18:1403-1412.
- Werner, R. A. 1978. The spruce beetle in Alaska forests. United States Department of Agriculture, Forest Service, Pacific Northwest Forest and Range Experiment Station. 8 p.
- Werner, R. A., R. D. Averill, F. L. Hastings, J. W. Hilgert, and U. E. Brady. 1984. Field evaluation of fenitrothion, permethrin, and chlorpyrifos for protecting white spruce trees from spruce beetle (Coleoptera: Scolytidae) attack in Alaska . Journal of Economic Entomology 77:995-998.
- Werner, R. A., B. H. Baker, and P. A. Rush. 1977. The spruce beetle in white spruce forests of Alaska. United States Department of Agriculture, Forest Service, Pacific Northwest Forest and Range Experiment Station, General Technical Report PNW 61. 13 p.
- Werner, R. A., E. E. Elert, and E. H. Holsten. 1983. Evaluation of beetle killed white spruce for pulp and paper . Canadian Journal of Forest Research 13:246-250.
- Werner, R. A., M. M. Furniss, L. C. Yarger, and T. Ward. 1983. Effects on eastern larch beetle of its natural attractant and synthetic pheromones in Alaska. United States Department of Agriculture, Forest Service, General Technical Report PNW 371. 7 p.
- Werner, R. A., F. L. Hasting, and R. D. Averill. 1983. Laboratory and field evaluation of insecticides against the spruce beetle (Coleoptera: Scolytidae) and parasites and predators in Alaska . Journal of Economic Entomology 76:1144-1147.
- Werner, R. A., and E. H. Holsten. 1983. Mortality of white spruce during a spruce beetle outbreak on the Kenai Peninsula in Alaska . Canadian Journal of Forest Research 13:96-101.
- Werner, R. A., and E. H. Holsten. 1984. Scolytidae associated with felled white spruce in Alaska . Canadian Entomologist 116:465-472.
- Holsten, E. H., P. E. Hennon, and R. A. Werner. 1985. Insects and diseases of Alaskan forests. United States Department of Agriculture, Forest Service, Alaska Region, Report 181. Juneau, Alaska. 217 p.
- Holsten, E. H., and R. A. Werner. 1985a. Evaluation of a controlled release formulation of methylcyclohexenone (MCH) in preventing spruce beetle attacks in Alaska. United States Department of Agriculture, Forest Service, Alaska Region, Anchorage, Alaska, State and Private Forestry, Report R10-9. 9 p.
- Holsten, E. H., and R. A. Werner. 1985b. Evaluation of a controlled release formulation of methylcyclohexenone (MCH) in preventing spruce beetle attacks in Alaska. United States Department of Agriculture, Forest Service, Alaska Region, Anchorage, Alaska, State and Private Forestry, Report R10-12. 9 p.
- Holsten, E. H., and R. A. Werner. 1987. Engraver beetles in Alaska forests. United States Department of Agriculture, Forest Service, Pacific Northwest Research Station, Portland, Oregon, undated circular, GPO 795-322. 6 p.
- Holsten, E. H., K. P. Zogas, R. L. Wolfe, and R. A. Werner. 1979. Resurrection Creek spruce beetle infestation: a three year interim report. United States Department of Agriculture, Forest Service, Alaska Region, Anchorage, Alaska, State and Private Forestry, Report.
- Miller, L. K., and R. A. Werner. 1987. Coldhardiness of adult and larval spruce beetles Dendroctonus rufipennis (Kirby) in interior Alaska . Canadian Journal of Zoology 65:2927-2930.
- Miller, M. C., J. C. Moser, M. D. McGregor, J. C. Gregoire, M. Baisier, D. L. Dahlsten, and R. A. Werner. 1987. Potential for biological control of native North American Dendroctonus beetles (Coleoptera: Scolytidae) . Entomological Society of America, Annals 80:417-428.
- Werner, R. A. 1986a. The eastern larch beetle in Alaska. United States Department of Agriculture, Forest Service, Pacific Northwest Forest and Range Experiment Station, Portland, Oregon, Research Paper PNW-357. ii + 13 p.
- Werner, R. A. 1986b. Association of plants and phytophagous insects in taiga forest ecosystems. Pages 205-212 in K. Van Cleve, F. S. Chapin III, P. W Flanagan, et al., Forest ecosystems in the Alaskan taiga: a synthesis of structure and function. Springer, New York.
- Werner, R. A. 1988a. Recommendations for suppression of an Ips perturbatus outbreak in interior Alaska using integrated control. Pages 189-195 in T. L. Payne and H. Saarenmaa, Integrated control of scolytid bark beetles. Virginia Polytechnic and State University, Blacksburg, Virginia. 355 p.
- Werner, R. A. 1988b. Suppression of an Ips perturbatus outbreak in interior Alaska using integrated control . International Congress of Entomology, Proceedings 18:414.
- Werner, R. A., J. S. Hard, and E. H. Holsten. 1988. The development of management strategies to reduce the impact of the spruce beetle in south-central Alaska . Northwest Environmental Journal 4:319-358.
- Werner, R. A., F. L. Hastings, E. H. Holsten, and A. S. Jones. 1986. Carbaryl and lindane protect white spruce from attack by spruce beetles (Coleoptera: Scolytidae) for three growing seasons . Journal of Economic Entomology 79:1121-1124.
- Werner, R. A., and E. H. Holsten. 1985a. Effect of phloem temperature on development of spruce beetles in Alaska. Pages 155-163 in L. Safranyik, The role of the host in the population dynamics of forest insects. IUFRO conference proceedings, Canada Department of the Environment, Forestry Service, and United States Department of Agriculture, Forest Service, Banff, Alberta, Canada, September, 1983. 240 p.
- Werner, R. A., and E. H. Holsten. 1985b. Factors influencing generation times of spruce beetles in Alaska . Canadian Journal of Forest Research 15:438-443.
- Werner, R. A., E. H. Holsten, and F. L. Hastings. 1986. Evaluation of pine oil for protecting white spruce from spruce beetle (Coleoptera: Scolytidae) attack . Entomological Society of British Columbia, Journal 83:3-5.
- Dixon, E., H. Wieser, H. Cerezke, E. Begin, G. Gries, R. Werner, and A. Whitehead. 1992. 1-methylcyclohex-2-enol, 3-methyl-cyclohex-22-enol and 3-methyl-cyclohex-2-enone as pheromones in three species of Dendroctonus bark beetles. Page 171 In Douglas C. Allen and L.P. Abrahamson (eds.), Proceedings: North American Forest Insect Work Conference, Denver, Colorado, March 25-28, 1991. United States Department of Agriculture, Forest Service, Pacific Northwest Research Station (Portland, OR), General Technical Report PNW-GTR-294.
- Holsten, E., and R. Werner. 1990. Comparison of white, sitka and Lutz spruce as hosts of the spruce beetle in Alaska . Canadian Journal of Forest Research 20:292-297.
- Holsten, E., and R. Werner. 1993. Effectiveness of polyethylene sheeting in controlling spruce beetles (Coleoptera: Scolytidae) in infested stacks of spruce firewood in Alaska. United States Department of Agriculture, Forest Service, Pacific Northwest Research Station (Portland, OR), Research Paper PNW-RP-466.
- Illman, B., and R. Werner. 1994. Sitka spruce response to Leptographium abietinum . Phytopathology 84:1143.
- Werner, R. 1993. Response of the engraver beetle, Ips perturbatus to semiochemicals in white spruce stands of interior Alaska. United States Department of Agriculture, Forest Service, Pacific Northwest Research Station (Portland, OR), Research Paper PNW-RP-465.
- Werner, R. 1994a. Research on the use of semiochemicals to manage spruce beetles in Alaska. Pages 15-21 In Patrick J. Shea (tech. coord.), Proceedings of the Symposium on Management of Western Bark Beetles with Pheromones: Research and Development, Kailua-Kona, Hawaii, June 22-25, 1992. United States Department of Agriculture, Forest Service, Pacific Southwest Research Station (Albany, CA), General Technical Report PSW-GTR-150.
- Werner, R. 1994b. Forest insect research in boreal forests of Alaska . Entomology 2:35-46.
- Werner, R., and J. Hilgert. 1992. Effects of permethrin on aquatic organisms in a freshwater stream in south-central Alaska . Journal of Economic Entomology 85:860-864.
- Werner, R., and E. Holsten. 1992. Effectiveness of sevin with or without diesel for remedial control of the spruce beetle (Coleoptera: Scolytidae) in infested spruce in Alaska . Journal of Economic Entomology 85:473-476.
- Werner, R., and B. Illman. 1994. Response of Lutz, Sitka and white spruce to attack by Dendroctonus rufipennis (Coleoptera: Scolytidae) and blue stain fungi . Environmental Entomology 23:472-478.
- Zhong, H., F. Hastings, F. Hain, and R. Werner. 1994. Toxicity of carbaryl toward the southern pine beetle in filter paper, bark and cut bolt bioassays . Journal of Entomological Science 29:247-253.
- Borden, J., G. Gries, L. Chong, R. Werner, E. Holsten, H. Wieser, E. Dixon, and H. Cerezke. 1996. Regionally-specific bioactivity of two new pheromones for Dendroctonus rufipennis (Kirby) (Col, Scolytidae) . Journal of Applied Entomology 120:321-326.
- Gara, R., R. Werner, M. Whitmore, and E. Holsten. 1995. Arthropod associates of the spruce beetle Dendroctonus rufipennis (Kirby) (Col, Scolytidae) in spruce stands of south- central and interior Alaska . Journal of Applied Entomology 119:585-590.
- Holsten, E., R. Werner, and R. Develice. 1995. Effects of a spruce beetle (Coleoptera: Scolytidae) outbreak and fire on Lutz spruce in Alaska . Environmental Entomology 24:1539-1547.
- McCuUough, D., R. Werner, and D. Neumann. 1998. Fire and insects in northern and boreal forest ecosystems of North America . Annual Review of Entomology 43:107-127.
- Werner, R. 1995. Toxicity and repellency of 4-allylanisole and monoterpenes from white spruce and tamarack to the spruce and eastern larch beetle (Coleoptera: Scolytidae) . Environmental Entomology 24:372-379.
- Werner, R. 1999. Population levels of bark beetles and associated insects in managed and unmanaged spruce stands . Forest Ecology and Management 115:267-275.
- Werner, R., and E. Holsten. 1995. Current status of research with the spruce beetle, Dendroctonus rufipennis. Pages 23-29 In S.M. Salom and K.R. Hobson (tech. eds.), Application of semiochemicals for management of bark beetle infestations — Proceedings of an informal conference. United States Department of Agriculture, Forest Service, Intermountain Research Station (Ogden, UT), General Technical Report INT-GTR-318. 54 pp.
- Werner, R., and E. Holsten. 1997. Dispersal of the spruce beetle, Dendroctonus rufipennis, and the engraver beetle, Ips perturbatus, in Alaska. United States Department of Agriculture, Forest Service, Pacific Northwest Research Station (Portland, OR), Research Paper PNW-RP-501. 8 pp.